Little Green
Sedge
Carex viridula
Sedge family (Cyperaceae)
Description:
This sedge consists of a tuft of leafy culms about ½–1½' tall; both
fertile shoots and infertile shoots are present. The
culms are light green, 3-angled, and glabrous. About 1-3 alternate
leaves occur along each culm; their blades are ascending to spreading
and arching. Individual leaf blades are up to 6" long and 3.5 mm.
across; they are light to medium green, glabrous, and centrally
grooved. The two outer sides of leaf sheaths are light to medium green,
glabrous, and vertically veined; the inner side of leaf sheaths is
membranous and glabrous. The inner leaf sheaths are concave to truncate
at their mouths; their short-membranous ligules are wider than they are
long. Each fertile culm terminates in an inflorescence consisting of
2-6 pistillate (female) spikelets and a terminal staminate (male) or
gynecandrous (female above and male below) spikelet. These spikelets
are usually clustered together, although the lower pistillate spikelets
are sometimes remote from the other spikelets. These spikelets are
sessile to short-pedunculate (without basal stalks or with short basal
stalks).
The pistillate spikelets are 5-12 mm. long, 4-5 mm. across,
and light green to yellow; they are short-cylindrical to nearly globoid
in
shape and prickly in appearance because of the short beaks of the
perigynia pointing in all directions. At the bases of lower pistillate
spikelets or their peduncles (basal stalks), there are solitary leafy
bracts up to 5" long and 3 mm. across; they are similar to the
alternate leaves below. The terminal spikelet, when it is staminate, is
about 6-8 mm. long, yellowish brown to brown, and narrow; when the
terminate spikelet is gynecandrous, it is about the same length,
although wider where the perigynia occur. The perigynia (sacs
surrounding the achenes) are 2-3 mm. long and about 1.5 mm. across;
they
are light green to yellow, glabrous, vertically veined, bluntly
3-angled, and ovoid-obovoid to obovoid in shape with a narrow straight
beak (about 0.5 mm. in length) above. The pistillate scales are a
little shorter than the perigynia and ovate-obovate to obovate in
shape; they are green-veined in the middle, otherwise membranous,
becoming brown or reddish brown with age. The blooming period can occur
from late spring to mid-summer, lasting 1-2 weeks for each plant.
Cross-pollination of the florets is by wind. Afterwards, the achenes
become mature about 1 month later. Mature achenes are 1.0–1.25 mm.
long, 0.75–1.0 across, obovoid and bluntly 3-angled in shape, glabrous,
and brown to black. The root system is fibrous and short-rhizomatous.
Cultivation:
The preference is full or partial sun, wet to moist conditions, and a
rather barren soil containing sand or gravel. Because this is a boreal
sedge, it is not well-adapted to hot dry weather.
Range &
Habitat: Little Green Sedge occurs in NE
Illinois, where it is native,
rare, and state-listed as 'threatened' (see Distribution
Map).
Illinois
lies along the southern range-limit of this species. This sedge has a
wide distribution in boreal areas of North America; it also occurs in
Eurasia. Only the typical subspecies of this sedge, Carex
viridula
viridula, is found in Illinois. Habitats include sandy shores
of small
ponds, interdunal swales along Lake Michigan, pannes (moist sand flats)
along Lake Michigan, seeps, fens, sandy marshes, and poorly drained
flat gravelly areas. In Illinois, this sedge is found in high quality
natural areas.
Faunal
Associations: Insects that feed on wetland
sedges (Carex spp.) include
leaf beetles
(Plateumaris spp.), billbugs (Sphenophorus
spp.), larvae of leaf-miner
flies (Cerodontha spp.), seed bugs (Cymus
angustatus, Oedancala
dorsalis), plant bugs (Mimoceps insignis, Teratocoris
discolor), aphids
(Allaphis spp., Iziphya spp., Rhopalosiphum
spp., Subsaltuaphis spp.),
leafhoppers (Cosmotettix spp.), sedge grasshoppers (Stethophyma
spp.),
larvae of the Tufted Sedge Moth (Hypocoena inquinata)
and other moths,
larvae of skippers (Euphyes spp., Poanes
spp.), and larvae of two
butterflies, Appalachian Brown (Satyrodes appalachia)
and Eyed Brown
(Satyrodes eurydice); see Clark et al. (2004),
Vaurie (1983), Spencer
& Steyskal (1986), Hoffman (1996), Marshall (2006), Knight
(1941),
Hottes & Frison (1931), Blackman & Eastop (2013),
Panzer et al.
(2006), Capinera et al. (2004), and Bouseman et al. (2006). An aphid,
Iziphya americana, has been observed to feed on
the Little Green Sedge
(Carex viridula). The seeds or seedheads of wetland
sedges are a source
of food for many birds, including ducks, rails, some songbirds,
and others. The Bird Table lists
some of these species. The Canada
Goose
also feeds on the leaves of these plants. White-tailed Deer browse on
the leaves and seedheads to a limited extent – the seeds
of sedges are able to pass through the digestive tract of this animal
and remain viable (Myers et al., 2004). This helps to spread the seeds
to new locations. Muskrats feed on the rhizomes and culms of sedges
to a limited extent (Hamerstrom & Blake, 1939), while some
turtles,
like the Snapping Turtle and Easter Mud Turtle, eat the seedheads
sparingly (Ernst et al., 1994).
Photographic
Location: Sandy shore of a small pond and a
moist sand flat along Lake
Michigan at the Illinois Beach State Park in NE Illinois.
Comments:
The Little Green Sedge (Carex viridula) is one of
the smaller wetland
sedges. Its leaf blades are up to 3.5 mm. across, its perigynia are 2-3
mm. long, and the beaks of its perigynia are about one-fourth as long
as their bodies. It is most similar to two other sedges that are found
in NE Illinois; they are also rare. These two species are Northeastern
Sedge (Carex cryptolepis) and Yellow Sedge (Carex
flava). Compared to
the Little Green Sedge, Northeastern Sedge has larger perigynia
(3.0–4.5 mm. long), and the beaks of its perigynia are proportionately
longer (about one-half the length of their bodies); the leaf blades of
this sedge also tend to be slightly wider (up to 4.5 mm. across).
Similarly, Yellow Sedge has even larger perigynia (4.0–6.0 mm. long),
and the beaks of its perigynia are also proportionately longer (about
the same length as their bodies). In addition, the leaf blades of this
last sedge are substantially wider (up to 6.5 mm. across) than those of
the Little Green Sedge. Another species, Hale's Sedge (Carex
haleana),
has perigynia that are similar in size to those of the Little
Green Sedge, but they lack significant beaks. As a result, the female
(pistillate) spikelets of Hale's Sedge have a grainy appearance, rather
than a prickly one. This sedge also has much wider leaf blades (up to
10 mm. across). A discussion of these and other technical differences
between these sedges can be found in Mohlenbrock (2011), especially on
p. 411.