Description:
This
perennial grass is 2-4' tall at maturity. It forms a stiff
culm about 2-30" tall that is terete and glabrous; this culm remains
quite short if an inflorescence is not
produced, in which case it is largely covered by the sheaths. Alternate
leaves occur primarily toward the base of
the culm. The leaf sheaths are light to medium green,
longitudinally veined, and glabrous; dried brown remnants of older
sheaths often persist at the base of the culm. The leaf blades are
up to 2½' long and 4-10 mm. across; they are ascending, stiff, and
straight. The lower half of each blade toward its base is usually flat,
while the upper half toward its tip is often strongly rolled
(involute). The inner surface of the blade (facing toward the culm) is
dull pale blue, while the outer surface of the blade (facing away
from the culm) is medium to dark green and glabrous. The margins
are rough-textured toward the tip of the blade, becoming more smooth
toward the base of the blade. The ligules are short-membranous and
about 3 mm. (1/8") long.
Occasionally, a culm terminates in a
spike-like panicle about 6-14" long and less than ¾" across. Each
panicle has a central stalk (rachis) and several lateral branches up to
2" long that are erect. Several single-flowered
spikelets on short pedicels occur along each lateral branch of the
panicle. Individual spikelets are 9-15 mm. long and slightly
flattened, consisting of a pair of glumes, a lemma, a membranous palea,
and a single floret. The glumes are 9-15 mm. long, linear-lanceolate
in shape and keeled along their outer surfaces. The lemma is a little
shorter than the glumes and linear-oblong in shape; there is a small
tuft
of hairs at its base. The floret consists of an ovary, 3 stamens, and a
pair of stigmata. The blooming period can occur from late spring to
late summer. The florets are wind-pollinated. Afterwards, fertile
lemmas are
replaced by elongated grains about 3.0-3.5 mm. in length. The root
system is fibrous and
long-rhizomatous. Reproduction occurs primarily by the rhizomes, which
can form large colonies of plants.
Cultivation:
The
preference is full sun, mesic to dry conditions, and very sandy
soil. Propagation occurs primarily by dividing clumps of plants and
their rhizomes, as most seeds are not viable. This grass can withstand
burial by sand up to 1' in depth. The root system is dependent on the
presence of mycorrhizal fungi to extract nutrients from the very sandy
soil where this grass occurs. Away from beaches and neighboring sand
dunes, this grass succumbs to damage from nematodes, making its
establishment more difficult, if not impossible.
Range
&
Habitat: The native Beach Grass is rare in Illinois and
state-listed as
'endangered.' It is found only in the NE section of the state along
Lake Michigan (see
Distribution
Map).
In the United States, this grass occurs along beaches of the Great
Lakes and the Atlantic coast. Habitats are restricted to drier areas of
sandy beaches
and unstable or partially stable sand dunes along Lake Michigan
and the other Great Lakes. This grass also occurs along the Atlantic
coast of northeastern United States. It plays an important role in
stabilizing sand in these habitats and preventing blowouts. Eventually,
as organic matter builds up, this sets the stage for the colonization
of other species of plants. In addition to destruction of habitat by
urban development, Beach Grass is vulnerable to trampling by
pedestrians and the use of off-road vehicles. Where the
latter problems exist, it can be protected by
erecting fences around colonies of plants.
Faunal Associations: The following information has been
derived
from published
material about Beach Grass populations along both the Atlantic coast
and Great Lakes. Some faunal species may not occur in both areas. The
following insects have been observed to feed on Beach Grass:
Eriococcus
carolinae (Beach Grass Scale), which sucks sap from the
stems and
leaves; larvae of the gall midge
Mayetiola
ammophilae, which feed
within the stem bases, causing them to rot;
Strigoderma arboricola
(False Japanese Beetle), which feeds on the florets and developing
seeds; a June Beetle,
Phyllophaga
rugosa; larvae of the Gelechiid
moth,
Chionodes arenella;
and
Trimerotropis
verruculata (Crackling
Forest Grasshopper), which sometimes feeds on the leaves. The
root system of Beach Grass is attacked by
Meloidogyne sasseri
(Beach
Grass Root-Knot Nematode) and other nematodes. Birds and other
vertebrate animals use Beach Grass primarily for cover and nesting
habitat; examples include the Short-Eared Owl, Piping Plover, Vesper
Sparrow, and various gulls. The Snow Bunting and probably other
granivorous birds eat the seeds to a minor extent. For more information
about floral-faunal relationships for this grass, see Eastman (2003)
and Maun (2009).
Photographic
Location: A large sand dune along Lake Michigan at the
Indiana Dunes State Park in NW Indiana.
Comments:
Although its population has declined, particularly in Cook County, this
is the dominant grass of beaches and sand dunes along southern Lake
Michigan. The stiff ascending leaves provide Beach Grass with a unique
appearance, making it easy to identify. The large inflorescence is also
rather showy. Another species in this genus, the non-native
Ammophila
arenaria (European Beach Grass), differs by having much
longer ligules.
This latter species is not found in Illinois, although it has been
introduced successfully along the Pacific coast in northwestern United
States.